Probably because of this bug https://jira.atlassian.com/browse/JSD-1792. Check if the information in the bug report relate to the problem you are seeing in your JIRA instance.
Cheers!
Teilor
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When it's ready and Atlassian have replicated and fixed it. Watch (and vote) on the issue to see the progress.
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Coudl you tell us more about the actual error? Is it just an NPE, or is there more error messages around it?
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The error we are seeing in the mail audit log is: Exception when MailHandlerWorker handles message: null
IRA MAIL Handler JIRA MAIL Handler[10021]: Cannot handle message as Issue Type is null or invalid. 2015-04-25 10:47:54,812 WARN [JEMH] QuartzWorker-0 ServiceRunner JIRA MAIL Handler JIRA MAIL Handler[10021]: Cannot handle message as Issue Type is null or invalid. 2015-04-25 10:48:05,652 ERROR [JEMH] QuartzWorker-0 ServiceRunner JIRA MAIL Handler JIRA MAIL Handler[10021]: Exception: javax.mail.MessagingException: Exception reading response; nested exception is:
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In this case I believe you are facing the know problem described in this KB: https://confluence.atlassian.com/pages/viewpage.action?pageId=224398784
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